NELSON AND GOLDMAN-REVISION OF JAGUARS 
 
regarded skulls from the valley of the Rio Tocantins as the most likely,
among 
those at hand, to represent this form, and have used them as a basis for

comparisons. 
Measurements.-No external measurements available. Skull (see table, p. 232).

Specimens examined.-Three, from Brazil, as follows: 
PARA: Lower Amazon, 1 (skull only'); Tocantins River Valley, 2 (skulls only).

Felis onca coxi, subsp. nov. 
Espiritu Santo Jaguar 
Type.-From north of Rio Doce, Espiritu Santo, Brazil. No. 256388 [9 ad.],

skull only, U. S. National Museum, collected by W. T. Cox, 1931. 
Distribution.-Eastern Minas Geraes and Espiritu Santo, Brazil; limits of

range unknown. 
General characters.-A small, light buffy or straw-colored subspecies. Much

smaller than its geographic neighbors, typical onca or the Matto Grosso animal,

described beyond, resembling the latter in color but differing from both
in cranial 
details. 
Color.-Skin from upper Rio Doce, Minas Geraes: Ground color of upper parts

in general light ochraceous buff or yellowish straw color, slightly darker
within 
the rosettes than in the interspaces, becoming still paler, less yellowish,
and near 
light buff on sides of neck, flanks, and outer surfaces of legs; black spots
and 
rosettes rather large and heavy, some enclosing smaller spots as in the other
forms; 
black spots large, irregular, and partly confluent along median line on posterior

part of back; under parts and inner surfaces of legs white, heavily spotted
with 
black; tail with irregular heavy black markings throughout its length, separated

by narrow interspaces, light buffy near base above, becoming white thinly
mixed 
with black toward tip and white below. 
Skull.--Cranium small, with a low sagittal crest and well-developed lambdoid

crest. Compared with that of subspecies onca the skull is much smaller; frontal

region more constricted immediately behind postorbital processes; bullae
rela- 
tively broader, somewhat flatter posteriorly near line of contact with exoccipitals.

Very much smaller, less angular than that of F. o. milleri, the Matto Grosso

jaguar; frontal region much less highly arched, the sides most deeply constricted

immediately behind postorbital processes (sides of frontals inflated behind

postorbital processes in Matto Grosso form); fronto-parietal suture more
evenly 
transverse instead of forming a narrow V pointing forward on the median line.

Measurements.-Flat tanned skin from upper Rio Doce, Minas Geraes: Total 
length, 2190 mm.; tail, 565. Skull (see table, p. 232). 
Remarks.-F. o. coxi contrasts strongly in size with its much larger geo-

graphic neighbors in the Parana and Paraguay river valleys. Two skulls re-

ceived many years ago from Herman von Ihering, and probably taken in the

general vicinity of S~o Paulo, were at first assigned here but seem referable
to 
the form described on p. 228. This new subspecies is named for the collector

of the type, Mr. W. T. Cox, in recognition of his extensive studies of wild
life. 
Specimens examined.-Total number, 4, as follows: 
I Collection Amer. Mus. Nat. Hist.