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S iI iY lI GajiiAL OTAiY 
Kqay 1, 1935.                                         H. Clarke 
Morphology and Anatomy of Lygodium .iaponicum. 
The Sporangium. 
1. Development from a single marginal cell, appearing in 
acropetal succession. 
2. A monangial sorus. 
3. Terminal annulus. 
Gametophyte. 
1. Germinationl of the spore forms a protlallium at first 
unsymmetrical, older prot.llia are syrmmetrical, 
2. Sex organs resemble L;ios of ot~her leptosporangiate ferns. 
The Young Sporophyte. 
1. Juvenile leves. 
2. Intermediabe leaves. 
The Leaf. 
1. Adult leaves characterized by indeterminate growth, and by 
twining. 
2. Arrested growing points associated with the lcaflets. 
3. Fertile and sterilu piniac. 
L-  Stem. 
1. An exarch protostele. 
2. Protoxylem does not have spiral or annular elements. 
3. Tracheids are long, havc scalariform thickenings, and may 
be irregular in shape. 
4. Small amount of phloem present. 
The Root. 
1. Xylem of root is diarch. 
2. Storage roots are present as branchies from the adventitious 
roots. The storage roots have vevy small stele. 
Lite,'ature Cited 
Binford, R. The development of the Sporangium of Lygodium, Bot. Gaz. 
44:214. 1903. 
 
 

					
				
					
General Botany--May 1, 1935.        2.                 H. Clarke 
Boodle, L. A. Comparative anatomy of tie Hymenophyllaceae, Schizaea- 
ceac, and Gleichhniceae II. On the anatomy of the Schizaea- 
ceae. Annals of Botany   15:559-421. 190J, 
vwer, F. 0. Studies in the morpology of tlie spore-producing 
membors, IV. Leptosporaigiat  Fms.    Ann. Lot. 13:320- 
324. 1899. 
-- -------Studies in the phylogeny of the Filicales, VII. Aim i 
Bot0  32:1-68. 1018. 
The u.ormal appcndages of ferns. Am.. Bet. 40:482. 
The ferns. Cambrid, Univ. Press. Vol. JI:155-17., 
Britton & Taylor. Schizaea psilla. Bull. lorroy Bot. Club. 28:1 
Diels, In Engler -aid Prantl, Vol. I:359-Z6. 
Cwyanc-Vaughn, D. T. The petiole of LygodiuL. Ani. Bet. 30:178. 
1916. 
LoLm, Carl. Untrsuchungen ubur Farnprot lallium. Flora 82:329- 
371. 1896. 
Luhl, Margaret. The histolo'y of thie sirv; tubcs of Pteridium 
Acnilinum, with so-ri notes oi, k>rsil1ia quadrifolia  and 
Lygodiu dichotomuL. Ani. Lot. 26:57--588. 
Litirdiere, Rene de. a'clerchs sar l ILeLit cihronosomique dans la 
caryocirese souatic des ilicineas. Ls 2ollule 31. 1920. 
' rantl, K. Untersucnungen zur Morpliooji. der Gcf~sskryptcgaien. 
Leipzig, 1881. 
:rs, Lenette. Developnent of te pror-haliium in Lygodiu  oal mu m 
Bet. Gaz. 75:75-81. 1923. 
-----------Developcnt of t ie archegone and studies in the 
fertilization of Lygodium palatuL. La Cellule 36: 
327-352. 1926. 
Stephens, Wo. Chase. On the developiment of the s,     " and spores

or Aneiraia Ahyl.itidis. A. :ct. 25:1039-1069. 
Twiss, Edith iv. Th  prothaliia of Al i ,... axf' L7 g 'iur. Bot. Gaz. 
49 :168. 2110 .